The first lesson of this course noted in passing that a meridian and its anti-meridian together complete a full circle around the Earth. This lesson is where that fact grows into a subject: the geometry of circles on a sphere, and what it costs, in miles and in degrees, to fly one kind of line rather than the other.
Great circles and small circles
Cut a sphere with a plane and the cut is always a circle; which kind depends on where the plane passes:
a great circle is the intersection of the Earth's surface with a plane passing through the Earth's centre: it is the largest circle that can be drawn on the sphere, it divides the Earth into two equal halves, and through any two points that are not diametrically opposite each other runs exactly one great circle;
a small circle is the cut of any plane that does not pass through the centre: every parallel of latitude except the equator is one.
A great circle that is not the equator and not a meridian reaches its vertex at the highest latitude it attains, one in each hemisphere, and at the vertex its direction is exactly east-west, 090 or 270 true. The two vertices lie 180° of longitude apart, and between them the circle crosses the equator twice. The Earth's ready-made examples of great circles are the equator and every meridian taken together with its anti-meridian; the parallels, cut by planes north or south of the centre, are the standing examples of small circles.
The great circle's claim to fame is economy: the shorter arc of the great circle through two positions is the shortest distance between them on the Earth's surface. The reason is the geometry of the sphere itself: of all the circles through the two points, the great circle is the one with the largest radius, and the flattest curve between two points is the shortest path a curved surface allows. Stretch a string tight between two points on a globe and it settles onto the great circle by itself.
The rhumb line
The alternative is built for the compass rather than the fuel gauge. A rhumb line (loxodrome) is a line on the Earth's surface cutting all meridians at the same angle: a line of constant true direction. Held long enough away from the special cases, it spirals ever tighter toward the pole without ever quite reaching it. The meridians, the equator and the parallels of latitude are all rhumb lines, and of them only the equator and the meridians run along great circles, a meridian pairing with its anti-meridian to complete one. Everywhere else, a rhumb-line route is not the shortest distance between two positions: its price for the constant heading is extra miles.
Convergence
Why can the great circle not simply hold one direction too? Because the reference it is measured against will not stand still. Direction is measured from the meridians, the meridians converge toward the poles, and a great-circle track therefore cuts each successive meridian at a different angle: its true direction changes continuously along the route, the equator and the meridians themselves being the only exceptions. The total change between two positions is the Earth convergence, and the syllabus approximates it with one formula: